C. Leaf

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Leaves are the most important appendages. They:

  • Carry out photosynthesis

  • Help remove excess water from plant body
  • Develop from leaf primordium
  • Are dorsiventrally flattened lateral appendage of stem

  • Produced at nodal region

  • Are thin, expanded, and green due to photosynthetic pigments

  • Show exogenous origin
  • Axil shows axillary bud

  • Show limited growth (no apical bud or growing point)


1. Typical Leaf Structure

A typical leaf shows three main parts:

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i. Leaf Base (Hypopodium)

  • Definition: Point by which leaf attaches to stem
  • Features: Varies in different plants
    • May be pulvinus (swollen)

    • Sheathing or ligulate

  • Stipules:
    • Pair of lateral outgrowths in some plants
    • Stipulate leaf: With stipules
    • Exstipulate leaf: Without stipules
    • Normally green protective structure

ii. Petiole (Mesopodium)

  • Definition: Part connecting leaf lamina with leaf base
  • Types:
    • Petiolate leaf: With petiole
    • Sessile leaf: Without petiole
  • Functions:
    • Helps lamina get exposed to light
    • Helps in conduction

iii. Lamina (Epipodium)

  • Structure: Large, expanded, flat, green part of leaf
  • Functions: Photosynthesis, gaseous exchange, transpiration
  • Anatomical types:
    • Dorsiventral (common in Dicots)

    • Isobilateral(common in Monocots)
    • Centric or cylindrical: Both surfaces not distinctly different
  • Variation: Lamina varies greatly in shape, margin, and apex

 


2. Leaf Venation

Definition: Arrangement of veins and veinlets in leaf lamina

Functions of Veins:

  • Conduction of water and minerals
  • Conduction of food
  • Development of structural framework of lamina

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3. Types of Leaf (Based on Incision of Lamina)

 

Simple Leaf

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  • Entire lamina (not divided)

Compound Leaf

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  • Lamina divided into number of leaflets

Pinnately Compound

  • Leaflets present laterally on common axis called rachis
  • Rachis represents the midrib of the leaf

Palmately Compound

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  • All leaflets attached at tip of petiole

 


4. Modifications of Leaves

Leaves may undergo modifications to perform functions other than photosynthesis (transpiration, gaseous exchange, light perception for flowering).

Leaf Spines

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  • Modification types:
    • Entire leaf modified into spines (Opuntia)
    • Leaf margin becomes spiny (Agave)
    • Stipule modified into spine (Acacia)
  • Function: Check rate of transpiration and protect from grazing
  • Examples: Zizyphus, etc.

Leaf Tendril

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  • Structure: Leaf, leaflet, or other part modified into thin, green, wiry, coiled structure

  • Function: Provide additional support to weak stems for climbing
  • Example: Weak-stemmed plants

Leaf Hooks

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  • Structure: Terminal leaflets modified into stiff, curved, pointed hooks
  • Function: Cling to tree bark
  • Example:

    Bignonia unguis-cati(Cat's nail)

Phyllode

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  • Definition: Petiole becomes flat, green, and leaf-like
  • Structure: Modified petiole replaces normal leaf
  • Example:Acacia auriculoformis
    • Normal leaf is bipinnately compound and falls off soon
    • Petiole modifies into phyllode
  • Significance: Xerophytic adaptation

Leaf Modifications

Category of Modification Type of Leaf Modification Structural Features Primary Function Examples
LEAF SPINES Entire Leaf Modified into Spines Complete leaf structure modified into sharp spines Check rate of transpiration, protection from grazing Opuntia
Leaf Margin Becomes Spiny Margins of leaf develop spiny projections while maintaining leaf-like structure Check rate of transpiration, protection from grazing Agave
Stipule Modified into Spine Stipule structures at leaf base become hardened and pointed Check rate of transpiration, protection from grazing AcaciaZizyphus
LEAF TENDRILS Leaf Tendrils Leaf, leaflet, or other leaf part modified into thin, green, wiry, coiled structure Provide additional support to weak stems for climbing Weak-stemmed plants
LEAF HOOKS Leaf Hooks Terminal leaflets modified into stiff, curved, pointed hooks Cling to tree bark for support and attachment Bignonia unguis-cati (Cat's nail)
PHYLLODE Phyllode Petiole becomes flat, green, and leaf-like; modified petiole replaces normal leaf; normal bipinnately compound leaf falls off soon Photosynthesis, xerophytic adaptation, reduced transpiration Acacia auriculoformis

5. Phyllotaxy

Definition: Arrangement of leaves on stem and branches in specific manner

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Function: Enables leaves to get sufficient light

Types:

Type

Description

Example

Alternate

Single leaf from each node

Mango

Opposite Decussate

Pair of leaves from each node; consecutive pairs at right angle

Calotropis

Opposite Superposed

Pair of leaves from each node; consecutive pair arranged directly above

Jamun

Whorled

Many leaves from each node

Nerium