D. Inflorescence

Definition of Inflorescence: A specialized axis or branch over which flowers are produced or borne in definite manner

Structure Parts:

  • Peduncle: Main axis
  • Flowers: Borne on peduncle

Significance

  • Not all flowers mature at same time
  • Advantages:
    • Increases chances of pollination
    • Large number of flowers can be pollinated in single visit
    • Makes plant attractive

Types of Inflorescence

Racemose Type

1

  • Growth of peduncle: Infinite or unlimited
  • Apical bud: Free for continuous growth
  • Flower arrangement:
    • Older flowers at base
    • Borne in acropetal succession
    • Order of opening is centripetal when peduncle is disc-shaped

Cymose Type

1

  • Growth of peduncle: Finite or limited
  • Apical meristem: Terminates into flower
  • Flower arrangement:
    • Oldest flower at apex
    • Borne in basipetal succession
    • Order of opening is centrifugal if peduncle is compressed from top

E. Flower

Flower: A highly modified and condensed shoot meant forsexual reproduction

1

Position A flower can be:

  • Axillary

    (borne in leaf axil)

  • Terminal(at tip of main axis)

Structure

Thalamus (Receptacle)

  • Consists of four compactly arranged nodes and three highly condensed internodes

  • From each node, a circle or whorl of modified leaves is produced

  • Color: Green (can perform photosynthesis)

Floral Terminology

Term

Definition

Example

Bracteate flower

Presence of bract at base of pedicel or over pedicel

-

Ebracteate flower

Absence of bract

-

Pedicellate flower

With pedicel (stalk)

-

Sessile flower

Without pedicel

-

Complete flower

Presence of all four floral whorls

-

Incomplete flower

Absence of any one floral whorl

-

Perfect/Bisexual/Monoclinous

Both androecium and gynoecium present

-

Imperfect/Unisexual/Diclinous

Any one reproductive whorl present

-

Staminate flower

Male flower only

-

Pistillate flower

Female flower only

-

Neuter flower

Both reproductive whorls absent

Ray florets of Tagetus (marigold)

Monoecious plant

Male and female flowers on same plant

Maize

Dioecious plant

Only one type of unisexual flowers on plant

Date palm, papaya

Zygomorphic(Bilateral symmetry)

Only one plane cuts flower showing mirror images

Sweet pea

Actinomorphic(Radial symmetry)

Can be cut along any diameter showing mirror images

Hibiscus

 


Insertion of Floral Whorls

Definition: Position and arrangement of floral whorls with respect to gynoecium on thalamus

Hypogyny

  • Ovary position: Highest position
  • Other parts: Below ovary
  • Ovary nature: Superior
  • Flower type: Hypogynous
  • Floral formula symbol:

     

  • Examples: Brinjal, Mustard, China rose
Perigyny
  • Positions: Ovary and other floral parts occupy about same position
  • Thalamus shape: Cup-shaped or saucer-shaped
  • Ovary nature: Semi-superior or semi-inferior
  • Arrangement: All floral whorls at rim of thalamus
  • Flower type: Perigynous
  • Floral formula symbol:

     

  • Examples: Rose, Pea, Bean
Epigyny
  • Thalamus shape: Closed cup-like
  • Ovary enclosure: Completely encloses ovary (may fuse with ovary wall)
  • Other parts: Occupy superior position
  • Ovary nature: Inferior
  • Flower type: Epigynous
  • Floral formula symbol:

     

  • Examples: Sunflower, Guava

Floral Parts and Their Structure Thalamus (Torus/Receptacle)

  • Called torus or receptacle
  • Green in color (can perform photosynthesis)

Calyx (K) Structure:

  • Outermost floral whorl
  • Members: Sepals
  • Usually green in color (perform photosynthesis)

Types:

  • Gamosepalous: Sepals united (e.g., China rose)
  • Polysepalous: Sepals free (e.g., Brassica)
Function

:

  • Protect inner floral parts in bud condition
  • Modify into pappus (stiff, hairy structures) for wind dispersal (e.g., Tridax, Sonchus)

Corolla (C)

Structure:

  • Second floral whorl
  • Members: Petals (variously colored)

Types:

  • Polypetalous: Petals free (e.g., Rose)
  • Gamopetalous: Petals fused (e.g., Datura)

Function:

  • Attract different agencies for pollination

Perianth (P)

Definition: Calyx and corolla remain undifferentiated

  • Members: Tepals

Types:

  • Polyphyllous: Tepals free
  • Gamophyllous: Tepals fused

Sub-types:

  • Sepaloid perianth: Green tepals
  • Petaloid perianth: Brightly colored tepals (e.g., Lily)

Aestivation (Arrangement in Bud)

Definition: Mode of arrangement of sepals, petals, or tepals in flower bud with respect to members of same whorl

Types:

Valvate

  • Margins remain in contact or lie close
  • Do not overlap
  • Example: Calyx of Datura

Twisted (Contorted)

  • One margin of a member is overlapped
  • Other margin overlaps adjacent member
  • Example: Corolla of China rose

Imbricate

  • One member internal (overlapped at both margins)
  • One member external (completely outside)
  • Rest overlap and get overlapped
  • Example: Calyx of Cassia

Vexillary

  • Corolla butterfly-shaped (5 petals)
  • Vexillum: Outermost and largest
  • Wings: Two lateral petals
  • Keel: Two smallest inner, fused petals
  • Example: Corolla of Pisum sativum

Epicalyx

  • Additional whorl of sepal-like structures
  • Formed by bractioles outside calyx
  • Characteristic feature

    of familyMalvaceae

Androecium (A) Definition: Third floral whorl (male reproductive part)

  • Members: Stamens

Condition: If all stamens are free = polyandrous

Stamen Parts:

Anther(terminal, produces pollen grains)

  • Usually

    bilobed/tetralocular/tetrasporangiate(e.g., Datura)

  • Sometimes

    monothecous(e.g., Hibiscus)

Filament(stalk, bears anther)

Connective(connects two anther lobes to filament)

Cohesion(Fusion within same whorl):

Adelphy

  • Stamens united by filaments
  • Anthers free
Syngeny
  • Anthers united
  • Filaments free
  • Example: Sunflower

Synandrous

  • Fused by both filaments and anthers
  • Example: Cucurbita
Adhesion

(Fusion between different whorls): Epipetalous

  • Stamens united with petals
  • Example: Datura

Epiphyllous

  • United with tepals
  • Example: Lily

Gynoecium (G)

Definition: Innermost floral whorl (female reproductive part/pistil)

  • Members: Carpels

Types:

  • Syncarpous: Carpels fused
  • Apocarpous: Carpels free

Carpel Parts:

Stigma

  • Terminal part
  • Receives pollen grains

Style

  • Narrow structure
  • Connects ovary with stigma

Ovary

  • Basal, swollen, fertile part
  • Produces ovules on placenta

Placentation (Arrangement of Ovules on Placenta)

Marginal

  • Ovules placed on

    fused marginsof unilocular ovary

  • Examples: Pea, Bean

Axile

  • Ovules placed oncentral axis

    of multilocular ovary

  • Examples: China rose, Cotton
Parietal
  • Ovules placed on

    inner wallof unilocular ovary of multicarpellary syncarpous gynoecium

  • Examples: Papaya, Cucumber

Basal

  • Single ovuleat base of unilocular ovary
  • Examples: Sunflower, Rice, Wheat

Free Central

  • Ovules borne oncentral axis

    not attached to ovary wall

  • Examples: Argemone, Dianthus, Celosia